The actors were shooting for We Live in Time at Herne Hill, sharing plenty of laughs together on and off camera.
Dressed in denim jeans and a brown sherpa jacket, Florence looked super cosy as she and Andrew strolled around the park.
Nailing the countryside chic vibe, the British star, 27, donned a pair of chunky boots and kept her make-up natural, her short blonde locks styled in a half-up, half-down do.
Meanwhile, the Tick, Tick… Boom! actor, 39, looked every inch the leading man in a blazer and blue shirt, paired with chinos as he carried a tote bag.
You’d be forgiven for assuming the co-stars were lifelong friends as they chatted over takeaway coffee, before heading to a market.
Outside the stall, Flo cheekily put a piece of fruit to Andrew’s mouth for him to smell, before he handed her a yellow flower.
Details of the plot of the film remain a closely-guarded secret, but the flick has been described as a ‘funny, deeply moving and immersive love story’.
Fans had already been dying for Florence and Andrew to appear on screen together, having never worked together before but stealing all our hearts when they presented the screenplay sscars together.
Speaking to Jimmy Fallon recently, Don’t Worry Darling star Florence confirmed the rumours that she and the Spider-Man actor would become co-stars.
‘We are doing a movie together… don’t worry that’s not a lie. We are doing a movie together.’
Proving they’re already getting on like a house on fire, she shared: ‘I met him the day before in rehearsals and we instantly got along and I was taking the p**s out of him and he loved it and he was taking the p**s out of me and I loved it. It was great!’
According to Deadline, the flick will be directed by John Crowley, with a script written by Nick Payne.
Sherlock’s Benedict Cumberbatch also serves as an executive producer, while Leah Clarke, Adam Ackland, and Guy Heeley are producing.
We Live In Time is yet another major project for in-demand Florence, whose schedule is jam packed.
It’s been a busy year for the movie star, who’s recent roles have been in Netflix’s The Wonder, as well as A Good Person.
This summer, she can be seen in Christopher Nolan’s Oppenheimer, and Dunne Part Two in November.
She’s also set to start filming Marvel Studios’ Thunderbolts, reprising her role as Yelena Belova.
As for Andrew, the role comes fresh from his success in FX limited series Under the Banner of Heaven, which earned him an Emmy and SAG Award nomination.
Prior to that, he earned an Oscar nom for his portrayal of Jonathan Larson and, of course, recently reprised his iconic role as Peter Parker in Spider-Man: No Way Home.
Got a story?
If you’ve got a celebrity story, video or pictures get in touch with the Metro.co.uk entertainment team by emailing us celebtips@metro.co.uk, calling 020 3615 2145 or by visiting our Submit Stuff page – we’d love to hear from you.
MORE : Victoria Beckham gives fans nostalgic tour of her childhood bedroom in Essex, featuring ‘funny-looking’ dolls and lavish furnishings
MORE : Michelle Williams recalls ‘unnerving’ attention over first Oscar nomination for Brokeback Mountain
var notifyQ = function () { var i = 0, l = awaitingReady.length; for (i = 0; i
For all the latest Entertainment News Click Here
For the latest news and updates, follow us on Google News.